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Cryptolithus lace collar trilobite fossils from Swatara Gap, Pennsylvania
Cryptolithus lace collar trilobite fossils from Swatara Gap, Pennsylvania.

Cryptolithus Trilobites: Lace Collar Trilobite Fossils

Cryptolithus is a small Ordovician trilobite famous for its wide perforated cephalon, or "lace collar." This guide explains its anatomy, filter-feeding lifestyle, fossil locations, and examples from Swatara Gap and the Kope Formation.


Beautiful Cryptolithus trilobite fossil showing the lace collar from Swatara Gap, Pennsylvania
Beautiful Cryptolithus trilobite fossil showing the lace collar. Fossil from Swatara Gap, Pennsylvania.

Fast Facts about Cryptolithus Trilobites

Cryptolithus trilobite fossils from Swatara Gap, Pennsylvania
Cryptolithus trilobite fossils from Swatara Gap, Pennsylvania.


Name: Cryptolithus is pronounced "crypto-LITH-us." The name means "hidden stone", from roots meaning hidden and stone.

Common name: Lace collar trilobite. The small holes, or fenestrae, around the edge of the cephalon look like a delicate lace collar.

Taxonomy: Arthropoda - Trilobita - Asaphida - Trinucleioidea - Trinucleidae - Cryptolithus.

Species: Commonly referenced species include Cryptolithus bellulus and Cryptolithus tessellatus.

Age: Ordovician Period.

Distribution: North America, especially Upper Ordovician deposits such as the Martinsburg Formation and Kope Formation.

Body size: Small. Most Cryptolithus trilobites are less than one inch long, or about 25 mm.

Diet: Filter feeder. The perforated fringe around the cephalon likely helped filter food particles from disturbed sediment or water near the sea floor.

Physical appearance: Cryptolithus has a wide cephalon with many small openings and very long genal spines. In complete specimens, the spines can be longer than the body of the trilobite.






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Cryptolithus Trilobite Facts and Information

Cryptolithus tessellatus trilobite restoration of appendages showing rarely preserved soft body parts
Cryptolithus tessellatus trilobite restoration showing appendages and soft body parts that rarely preserve. This is Figure 20 from Raymond (1920), drawn by Dr. Elvira Wood from original specimens and photographs made by Professor Beecher.

Cryptolithus trilobites are a genus of small, blind Ordovician trilobites. They are often called "lace collar" trilobites because of the small openings, or fenestrae, around the edge of the cephalon. These openings are the most distinctive feature of the genus and are easy to see in well-preserved specimens.

The lace-like fringe was not just decorative. It likely functioned as part of a feeding system. The trilobite probably disturbed sediment on the sea floor and used the perforated cephalic fringe to filter small food particles. This made Cryptolithus a specialized filter feeder living close to the bottom of Ordovician seas.

In addition to the wide perforated cephalon, lace collar trilobites are known for their extremely long genal spines. These spines can extend beyond the length of the body from cephalon to pygidium, giving complete specimens a delicate and dramatic appearance.

Although Cryptolithus is a small trilobite, it can be one of the most beautiful Ordovician fossils when complete. Unfortunately, complete specimens are uncommon. Most finds are fragments, especially pieces of the cephalon or isolated parts of the perforated fringe.




Where to Find Cryptolithus Trilobite Fossils

Cryptolithus fossils occur in Upper Ordovician rocks of North America. Two of the most important fossil sources are the Martinsburg Formation in Pennsylvania and the Kope Formation in the Cincinnati region.

The world-famous specimens of Cryptolithus bellulus come from the Martinsburg Formation. One famous collecting locality was Swatara Gap in Lebanon County, Pennsylvania. Here, the fossils are often preserved as casts coated with a fine rusty iron-oxide powder. This limonite coating gives the trilobites their distinctive color and makes complete specimens especially attractive.

Cryptolithus tessellatus is common in the Kope Formation, an extensive Upper Ordovician formation exposed in parts of Ohio, Kentucky, and Indiana. Complete specimens are uncommon in the Kope Formation, but fragments can be abundant in fossil hash plates composed of crinoid fragments, brachiopods, bryozoans, and trilobite pieces.


Cryptolithus fragments in a Kope Formation fossil hash plate
Cryptolithus fragments in a Kope Formation fossil hash plate.

The image above and the image below show hash plate sections from the Kope Formation. The matrix is composed mostly of crinoid fragments, along with brachiopods, bryozoans, and pieces of Cryptolithus trilobites.


Kope Formation matrix with crinoids and Cryptolithus fragments
Kope Formation matrix with crinoids and Cryptolithus fragments.




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Trilobites: Common Trilobites of North America (A NatureGuide Book)
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Cryptolithus Fossil Examples




Cryptolithus bellulus from Swatara Gap

Cryptolithus bellulus trilobite fossil from Swatara Gap Pennsylvania
A beautiful Cryptolithus bellulus preserved in iron-oxide powder. Notice the lace collar and long genal spines.

Formation: Martinsburg Formation
Age: Upper Ordovician
Location: Swatara Gap, Lebanon County, Pennsylvania


Complete Cryptolithus bellulus trilobite fossil from Swatara Gap Pennsylvania
Another complete Cryptolithus bellulus trilobite fossil. The lace collar is less visible from this angle.

Formation: Martinsburg Formation
Age: Upper Ordovician
Location: Swatara Gap, Lebanon County, Pennsylvania


Cryptolithus cephalon fossil from Swatara Gap Pennsylvania
A cephalon of Cryptolithus from Swatara Gap.

Formation: Martinsburg Formation
Age: Upper Ordovician
Location: Swatara Gap, Lebanon County, Pennsylvania


Cryptolithus tessellatus from the Kope Formation

Cryptolithus tessellatus fragments from the Kope Formation
Part of a hash plate from the Kope Formation showing numerous crinoid fragments, pieces of bryozoans and brachiopods, and Cryptolithus tessellatus trilobite fragments.

Formation: Kope Formation
Age: Ordovician
Location: Ohio


References / Works Cited


Raymond, P. E. (1920). The Appendages, Anatomy, and Relationships of Trilobites. Memoirs of the Connecticut Academy of Arts and Sciences. Yale University Press. Project Gutenberg ebook.


FAQs: Cryptolithus Trilobite Frequently Asked Questions

Learn what Cryptolithus trilobites were, why they are called lace collar trilobites, where they are found, and how their unusual cephalon helped them feed.


  • What is Cryptolithus?
    Cryptolithus is a genus of small Ordovician trilobites known for a wide perforated cephalic fringe, often called a lace collar. These openings helped the animal filter food particles from sediment or water near the sea floor.
    See: Cryptolithus Overview
  • Why is Cryptolithus called the lace collar trilobite?
    Cryptolithus is called the lace collar trilobite because the broad fringe around its cephalon is covered with small openings called fenestrae. These holes give the head shield a delicate lace-like appearance.
    See: Fast Facts


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