Cryptolithus is a small Ordovician trilobite famous for its wide perforated cephalon, or "lace collar." This guide explains its anatomy, filter-feeding lifestyle, fossil locations, and examples from Swatara Gap and the Kope Formation.
Species:
Commonly referenced species include Cryptolithus bellulus and Cryptolithus tessellatus.
Age:
Ordovician Period.
Distribution:
North America, especially Upper Ordovician deposits such as the Martinsburg Formation and Kope Formation.
Body size:
Small. Most Cryptolithus trilobites are less than one inch long, or about 25 mm.
Diet:
Filter feeder. The perforated fringe around the cephalon likely helped filter food particles from disturbed sediment or water near the sea floor.
Physical appearance:Cryptolithus has a wide cephalon with many small openings and very long genal spines. In complete specimens, the spines can be longer than the body of the trilobite.
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Cryptolithus Trilobite Facts and Information
Cryptolithus tessellatus trilobite restoration showing appendages and soft body parts that rarely preserve. This is Figure 20 from Raymond (1920), drawn by Dr. Elvira Wood from original specimens and photographs made by Professor Beecher.
Cryptolithus trilobites are a genus of small, blind Ordovician trilobites. They are often called "lace collar" trilobites because of the small openings, or fenestrae, around the edge of the cephalon. These openings are the most distinctive feature of the genus and are easy to see in well-preserved specimens.
The lace-like fringe was not just decorative. It likely functioned as part of a feeding system. The trilobite probably disturbed sediment on the sea floor and used the perforated cephalic fringe to filter small food particles. This made Cryptolithus a specialized filter feeder living close to the bottom of Ordovician seas.
In addition to the wide perforated cephalon, lace collar trilobites are known for their extremely long genal spines. These spines can extend beyond the length of the body from cephalon to pygidium, giving complete specimens a delicate and dramatic appearance.
Although Cryptolithus is a small trilobite, it can be one of the most beautiful Ordovician fossils when complete. Unfortunately, complete specimens are uncommon. Most finds are fragments, especially pieces of the cephalon or isolated parts of the perforated fringe.
Where to Find Cryptolithus Trilobite Fossils
Cryptolithus fossils occur in Upper Ordovician rocks of North America. Two of the most important fossil sources are the Martinsburg Formation in Pennsylvania and the Kope Formation in the Cincinnati region.
The world-famous specimens of Cryptolithus bellulus come from the Martinsburg Formation. One famous collecting locality was Swatara Gap in Lebanon County, Pennsylvania. Here, the fossils are often preserved as casts coated with a fine rusty iron-oxide powder. This limonite coating gives the trilobites their distinctive color and makes complete specimens especially attractive.
Cryptolithus tessellatus is common in the Kope Formation, an extensive Upper Ordovician formation exposed in parts of Ohio, Kentucky, and Indiana. Complete specimens are uncommon in the Kope Formation, but fragments can be abundant in fossil hash plates composed of crinoid fragments, brachiopods, bryozoans, and trilobite pieces.
Cryptolithus fragments in a Kope Formation fossil hash plate.
The image above and the image below show hash plate sections from the Kope Formation. The matrix is composed mostly of crinoid fragments, along with brachiopods, bryozoans, and pieces of Cryptolithus trilobites.
Kope Formation matrix with crinoids and Cryptolithus fragments.
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Cryptolithus Fossil Examples
Cryptolithus bellulus from Swatara Gap
A beautiful Cryptolithus bellulus preserved in iron-oxide powder. Notice the lace collar and long genal spines.
Part of a hash plate from the Kope Formation showing numerous crinoid fragments, pieces of bryozoans and brachiopods, and Cryptolithus tessellatus trilobite fragments.
Raymond, P. E. (1920). The Appendages, Anatomy, and Relationships of Trilobites. Memoirs of the Connecticut Academy of Arts and Sciences. Yale University Press.
Project Gutenberg ebook.
Learn what Cryptolithus trilobites were, why they are called lace collar trilobites, where they are found, and how their unusual cephalon helped them feed.
What is Cryptolithus?
Cryptolithus is a genus of small Ordovician trilobites known for a wide perforated cephalic fringe, often called a lace collar. These openings helped the animal filter food particles from sediment or water near the sea floor.
See: Cryptolithus Overview
Why is Cryptolithus called the lace collar trilobite?
Cryptolithus is called the lace collar trilobite because the broad fringe around its cephalon is covered with small openings called fenestrae. These holes give the head shield a delicate lace-like appearance.
See: Fast Facts
When did Cryptolithus live?
Cryptolithus lived during the Ordovician Period, more than 440 million years ago, in shallow marine environments of ancient North America.
See: Fast Facts
What did Cryptolithus eat?
Cryptolithus was likely a filter feeder. It probably stirred up sediment and used the perforated fringe on its cephalon to filter small food particles near the sea floor.
See: Cryptolithus Overview
Where are Cryptolithus fossils found?
Cryptolithus fossils are found in Upper Ordovician rocks of North America. Famous examples of Cryptolithus bellulus come from the Martinsburg Formation at Swatara Gap in Pennsylvania, while Cryptolithus tessellatus is common as fragments in the Kope Formation of Ohio, Kentucky, and Indiana.
See: Fossil Locations
How big were Cryptolithus trilobites?
Cryptolithus trilobites were small, usually less than one inch, or about 25 millimeters, long. Their long genal spines can make complete specimens appear larger.
See: Fast Facts
What are common Cryptolithus species?
Two well-known species are Cryptolithus bellulus, famous from Swatara Gap, Pennsylvania, and Cryptolithus tessellatus, commonly found as fragments in the Kope Formation.
See: Fossil Examples
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