This Thursday evening (Sept. 29) from 7-8 pm (eastern) is the FOSSIL Project's 2nd Fall Webinar!
Bruce MacFadden is doing the next one on "Field Notes 101"
Go to FOSSIL Webinar Series Forum for directions on logging in to participate in the free webinar!
Triceratops Dinosaur Specimen on display at CMNH
Fast Facts about Triceratops Dinosaurs
Triceratops (pronunciation: "tri-cer-ra-tops") - The name means "Three Horned Face"
Taxonomy: Dinosauria (Dinosaur) - Ornithischia (Bird Hipped) - Genasauria (Beaked Herbivor) - Cerapoda (Uneven Enamel on Lower Teeth) - Marginocephalia (Fringed Head) - Ceratopsia (Horned Face) - Neoceratopsia (New Ceratopsia) - Ceratopsoidea - Ceratopsidae (Family Level)
Age: Late Cretaceous
Triceratops appeared around 68 million years ago and went extinct around 65 million years ago during the end Cretaceous mass extinction event.
Discovery: Othniel C. Marsh, 1887
The first described specimen comes from Colorado. The specimen (a set of horns and skull section) was sent to Marsh for study. He concluded the horns were from a prehistoric bison. After other discoveries a year later, Marsh realized it was a dinosaur, and called it "Triceratops'.
Distribution: Western North America:
Triceratops fossils are only found in on the island continent of Laramidia, Which is now present day Western North America. It shared this island continent with T. rex.
Body Size: 30 feet long, and 10 feet tall!
Triceratops was built like a tank. It could grow to around 30 feet long, 10 feet tall, and could reach a weight of 12 tons (that's about 160 people). The horned head was a full 1/3 of its entire body length.
Triceratops was a herbavore. They roamed in large herds and ate plants.
Introduction to Triceratops and other Ceratopsid Dinosaurs - Fossil Facts and Information
Protoceratops andrewsi specimen at the CMNH
It is one of the few Asian Ceratopsid Dinosaurs.
It is a small Ceratopsid (6 feet in length).
The Ceratopsids - Origins and Distribution
The Ceratopsidae Dinosaur family encompasses the quadrupedal herbivores with distinguishing facial horns and large frills.
This Dinosaur family includes the well known Triceratops, which is the first Ceratopsid discovered back in 1887.
There are numerous subfamilies and genera of Ceratopsid. They are classified by the shape and size of their horns, beaks, and frills.
Ceratopsids are restricted to North America and Asia (Triceratops is only found in Western North America). Primitive Ceratopsids have been found in Asia, indicating they first appeared there and then crossed over to North America on a land bridge. Once in North America, they rapidly diversified until their extinction at the end of the Cretaceous. Approximately 37 genera of Ceratopsids have been classified. Below is an image of a few genera of Ceratopsid skulls, showing the distinctly different horn and frill shapes and sizes.
An image of skulls from some members of the Ceratopsid Dinosaur family. Notice the wild variation in the frill, beak, and horns. This image is from the Natural History Museum of Utah.
By skinnylawyer from Los Angeles, California, USA [CC-BY-SA-2.0], via Wikimedia Commons
Variation in Frills and Horns of the same species
When Ceratopsids were being discovered in the 1900's, they were classified based on their frill, beak,
and horn shapes alone. No consideration was given to juvenile vs.
adult appearance and sexual dimorphism. This is probably due to
the fact that almost all fossil remains were fragmentary and
incomplete. As a result, countless species were erected.
Later as more and more fossil evidence was discovered, paleontologists realized the skull changes shape as the animal enters different stages of development. Basically, horns and frills were not fully developed until adulthood (Farlow 327).
A great example of this is from the baby chasmosaurine ceratopsid dinosaur discovered in 2013. Currie et al, in 2016, published a paper on this baby dinosaur and compared it to adult chasmosarus specimens (Currie et al, 2016). The baby chasmosarus' frill looks completely different than the adult frill.
The Triceratops saga... from 16 species down to 2 - The great species roundup.
A Triceratops Dinosaur skeleton on display
at the Field Museum of Natural History in Chicago
A good example of reducing the number of Ceratopsids is the Triceratops genera. Triceratops is the most commonly found
Dinosaur in the Hell Creek Formation of the western United States. The Triceratops genus once had up to 16 species
assigned to it based on all the different skull variations found. Once paleontologists realized the frills changed
during development, Triceratops was reanalyzed. In 1986 Ostrom and Wellnhofer reduced Triceratops down to just 1 species,
T. horridus. They figured all the skull differences were due to different stages of development and sexual dimorphism,
and there couldn't be numerous similar large animals all living in the same area.
A little later, Catherine Forester in 1996 researched the Triceratops species and concluded since T. horridus greatly outnumbered T. prorsus, they were two distinct species, and not one sexually dimorphic species. Finally, in 2009 Scannella and Fowler supported this view, and the saga was over. There were two Triceratops species... sort of... There's that whole Torosaurus issue...
Triceratops vs. Torosaurus - Triceratops is not Torosaurs
Comparison of the size of Torosaurus latus specimens.
"A, YPM 1831, a subadult, skull length >2.6 m, and B, ANSP 15192,
an early adult, skull length 1.8 m. The rostrum of YPM 1831 is
reconstructed and would probably have been slightly longer,
as in ANSP 15192." From: Nicholas R. Longrich*, Daniel J. Field,
2012 - PLOS one under CC2 license.
Torosaurus is a large ceratopsid found in the same formation as Triceratops.
It is larger, and the frill is distinctly different, being longer, with large holes near the top.
In 2010 Scannella and Horner published a paper demonstrating that Torosaurus was actually a late, rapid,
growth stage of Triceratops. This stirred all kinds of controversy. Paleontologists were taking sides, some
agreed, and some disagreed. Personally, I always had trouble with how two large holes could simply erode in
the frill bone. That would really transform the entire muscle attachment structures. Within the last few years,
many papers have been published refuting Scannella and Horner's results. In 2011 the ceratopsid dinosaur expert
Dr. Andy Farke published a paper showing Torosaurus is "a genus distinct from Triceratops and Nedoceratops." Recent papers
tend to support the separate genus, including a paper by Longrich and Field in 2012 . They studied various Torosaurus
and Triceratops skulls, and did a clustering analysis to arrange them in growth series. They found immature and mature
individuals of Torosaurus and Triceratops. Therefore, they cannot be the same animal. Triceratops and Torosaurus are
The image to the left is from Longrich and Field (2012). It shows an adult and a subadult Torosoarus.
Horns and Frills - They Weren't Made For Ramming!
Ceratopsid Dinosaur skull from Utah showing its
elaborate Horns and Frills" on display at the CMNH
Most of the Ceratopsids had huge frills, with the exceptions of Diceratops and Triceratops, whose frills
were comparatively reduced in size (Fastovsky 180). Increases in frill size during life directly affected
jaw muscle strength (the larger the frill, the larger/stronger the jaw muscle) (Fastovsky 186). The big holes
seen in some of the frills were probably filled with muscle attachments and or cartilage.
As for the horns, many people forget when they see a fossil Triceratops or other Ceratopsid Dinosaurs, the horns on the skull are the bone cores. These bone cores would have been covered by the actual horns, so they would have been longer in life.
Unfortunately, it is impossible to verify what the horns and frills were actually used for. However, using modern herding animals as an analogy, the frills and horns it is probable they were used for dominance, defense, and display, just like living antelopes and deer. Therefore, Ceratopsids may have used their horns within their own species for territorialism and mating rights. Puncture wounds have been commonly found in skulls and frills of Triceratops from other Triceratops, indicating some form of intraspecies combat (Farke, et al, 2009). A study showed that if two adult Triceratops rammed each other head on, their skulls would shatter. However, Ceratopsid horns were shaped different than any mammilian counterpart. Instead of ramming, they may have simply locked horns and fought. A study using models found it was physically possible for triceratops to engage in horn locking behavior (Farke, Andrew, A. 2004). They found 3 possible horn locking arrangements. Instead of ramming each other, they would lock horns and fight, sort of like how moose will lock horns when fighting.
It is interesting to note, that although it was likely that Triceratops used their horns for combat, a different genera, Centrosaurus (A ceratopsid with only 1 large nasal horn) was studied by Farke (Farke, et al, 2009) and was found not have injuries. This means Centrosaurs probably did not use their horns and frills for mating or territorial combat, but instead for display purposes, like a peacock. A firgure from Farke's paper is shown to the right. It shows incidence rates of showing incidence rates of lesions in (A) Triceratops and (B) Centrosaurus.
"Schematics of the skulls of (A) Triceratops and (B) Centrosaurus, showing
incidence rates of lesions (periosteal reactive bone and fracture calluses)
on each cranial element (number of abnormal elements / total number of elements)
. Not to scale." From: Andrew A. Farke, Ewan D. S. Wolff, Darren H. Tanke, 2009
- PLOS one under CC2 license.
Moral of the story, if Triceratops used them to fight, while Centrosaurus used them for display, then horns and frills were
probably used in a variety of different ways by different types of ceratopsids.
One use of the horns and frills by many ceratopsids was probably to ward off predators. Some ceratopsid fossils have even been found with healed T-rex bite marks, these are specimens that have successfully staved off a T-Rex attack. While many others have been found with unhealed bite marks, meaning they were T-Rex food. Because of these fossil finds, it is not unreasunable to assume the frill and horns were used to fend off the mighty T-Rex that lived in the Western U.S. in the late Cretaceous.
Whether horns and frills were used for horn locking, head bunting, display, defence, or all of the above, one thing is for certain. They came in all shapes and sizes, as the image below shows!
Ceratopsid Dinosaur horns and frills on display at the Carnegie Museum of Natural History.
Notice the first skull to the left (Pachyrhinosaurus sp.) has no horn at all, instead it has a thick bone knob.
Sketch of a Triceratops Tooth
Large bone beds of 100's of Centrosaur fossils have been found in the Midwest, indicating they traveled in large herds. No large bone beds of Triceratops have been found, so they may have traveled in smaller groups, rather than large herds. This is consistent with the fact that horn usage is more prevalent in herding animals, such as deer, gazelle, and buffalo.
Like all herding animals, Ceratopsids were herbivores. Their mouths were ideally suited to eating tough vegetation. These Dinosaurs had double rooted teeth that were arranged in groups called dental batteries. A dental battery is a unique tooth set-up where adjacent teeth were locked together in longitudinal rows and in vertical columns. With at least 3 teeth in each column, 1 was functional and 2 were "replacements." The teeth, which were continually replaced throughout Dinosaurs' lives, were positioned so vertically and were so sharp (self-sharpening in fact) that they could shear like scissors. During chewing, these dinosaurs' teeth really did not touch at all like humans' do; instead they slid past each other, in a slicing motion (Fastovsky 183). Because of these razor sharp teeth and very strong jaw muscles, they are often referred to as "Nature's first Cuisinarts!!" (Farlow 85). They could slice through the toughest material.
Scientists are somewhat unsure of their exact plant diet. They suspect, due to their relatively low browsing height, that they may have eaten such plants as angiosperms, ferns, and small conifers growing at that height. Due to their large and heavy skulls, they could not rise up on their hind legs, but these lawnmowers of the American west may have been able to knock over trees in order to gain access to the leaves.
There is some debate on whether or not triceratops ate meat. Although the beak and teeth have been shown to be ideally suited for chomping up tough plant material, it has been shown the teeth are also good at chomping up anything... including meat. However, there has been no evidence found that Triceratops or other Ceratopsids ate meat. With that said, modern bovine (cows) and deer have been documented eating small birds and animal carcasses, yet we still call these animals herbivores. So if a malnourished Triceratops stumbled upon a carcass, would it pass it up? Who knows? Triceratops and other Ceratopsids are however considered herbivores... they ate plants!
How fast could Triceratops Dinosaur Run? - Triceratops and Ceratopsid Locomotion
Skeletal reconstructions of Ceratopsid Dinosaurs in many museums (including the ones pictured in this article,
the Carnegie Museum, and the Chicago Filed Museum) show Triceratops with a sprawling lizard like posture.
These dinosaurs should have a more mammal like posture, with the front legs underneath the animal, and not
splayed out to the side. The lizard like posture comes from errors when mounting the rib and vertebra of
Evidence supporting the above statements come from analysis of Triceratops and other Ceratopsian trackways. Paul and Christiansen published a paper in 2000 in which they studied the pedal impressions in trackways and analyzed the placement and angles of the hands and feet. From this information, they determined they have an elephant like forelimb posture. A mammal like posture enables Ceratopsids to run MUCH faster than the reptile like posture. Paul and Christiansen then indicate the maximum running speed of the largest Ceratopsids to be "broadly similar to that of rhinos." By the way, The Black Rhinoceros can run at speeds around 55 km/hr, or 34 miles per hour! That's pretty darn fast!
The left Triceratops skeletal reconstruction from the Carnegie Museum of Natural History shows a sprawling, lizard like posture, which according to recent evidence is incorrect.
The right Triceratops model from the Field Museum in Chicago shows a more mammal like posture, which according to recent evidence is correct
"Skin impression of Centrosaurus (Monoclonius cutleri)"
Date: 1917, Author: Barnum Brown,
Source: http://digitallibrary.amnh.org/.../B037a10.pdf Public Domain
Fossilized Dinosaur skin impressions are VERY rare to find. They often dub Dinosaur fossils with skin as "Dino Mummies"
although they are not mummified. The actual skin is not preserved, but impressions of the skin in the once soft mud
that surrounded the animal fossilize.
Fortunately, for Triceratops lovers, a "Triceratops Mummy" named Lane was discovered in Wyoming in 2002. The Black Hills Institute of Geological Research carefully and painstakingly excavated the T. horridus. It is now on display at their museum with another almost completely intact T. horridus skeleton called Kelsey.
The Triceratops "Lane" has the most preserved skin than any other Dinosaur fossil found. There are areas where several feet of skin are preserved. The skin is still being researched, and a paper has not been published as of yet. In the mean time, the Black Hills Institute has a cast of the skin displayed.
The skin is very different than what many paleontologists thought. The underside of the animal is covered in scutes, similar to crocodiles. The rest of the dinosaur is covered in small scales, like other dinosaur skin, however, there are also irregular scale like shapes. Some are hexagon in shape and about the size of a fist and have small pore-like holes in the center. What these are is any ones guess right now, as it is currently being studied.
Some people have speculated that quills may have protruded from these holes. This "Quill" hypothesis stems from a relative of the Triceratops, Psittacosaurus from Asia. It is a distant relative that has quill like structures on the top of its tail. It is important to note that Psittacosaurur is a distant Triceratops relative, so it does not mean Triceratops would have had quills also. The "Quill" hypothesis currently lies in the realm of speculation, and we will need to wait for further studies and publications shed light on the exact nature of Triceratops skin.
The skin impression to the right is not from "Lane," as all the Lane skin images are currently privately owned. It is one of the original skin impressions of a Ceratopsid found. It's from a Centrosaurus.
Triceratops had rows of teeth arranged in large "batteries" These teeth were continuously replaced. As a result, Triceratops teeth are fairly common. They make nice affordable gifts for dinosaur enthusiasts. Fossil Era has a nice selection of real Triceratops teeth.
The following book: New Perspectives on Horned Dinosaurs: The Royal Tyrrell Museum Ceratopsian Symposium (Life of the Past) gives an updated view on ceratopsians. It presents new research on the functions, biology, behavior, paleoecology, and paleogeography of horned dinosaurs. It also describes many new species. The book is a collection of scientific papers, so this is for serious dinosaur readers only! I think it's a great reference book and a must resource for dinosaur enthusiasts.
THE CONCISE DINOSAUR ENCYCLOPEDIA [ Hardcover ]
by Burnie, David. March 2004
This is one of the better general books on dinosaurs. It is incredibly visual and goes over countless dinosaurs lifestyles, behaviors, and the habitat they lived in. It also covers dinsoaur evolution and their extinction.
Currie Philip J., Holmes Robert B., J. Ryan Michael, Coy Clive. (2016) A juvenile chasmosaurine ceratopsid (Dinosauria, Ornithischia) from the Dinosaur Park Formation, Alberta, Canada. Journal of Vertebrate Paleontology, 2016; e1048348 DOI: 10.1080/02724634.2015.1048348
Farlow, O. James, Brett-Surman, M.K., Editors (1997). The Complete Dinosaur. IN: Indiana University Press.
Fastovsky, E. David, Weishampel, B. David (1996). The Evolution And Extinction Of The Dinosaurs. NY: Cambridge University Press.
Farke A.A. (2011) Anatomy and Taxonomic Status of the Chasmosaurine Ceratopsid Nedoceratops hatcheri from the Upper Cretaceous Lance Formation of Wyoming, U.S.A. PLoS ONE 6(1): e16196. doi: 10.1371/journal.pone.0016196
Farke AA, Wolff EDS, Tanke DH. (2009) Evidence of Combat in Triceratops. PLoS ONE 4(1): e4252. doi: 10.1371/journal.pone.0004252
Farke, A.A. 2004. Horn use in Triceratops (Dinosauria: Ceratopsidae): Testing behavioral hypotheses using scale models. Palaeontologia Electronica 7(1):10 pp. (PDF)
Forster Catherine A. (1996) Species resolution in Triceratops: cladistic and morphometric approaches. Journal of Vertebrate Paleontology, 16:259-270.
Longrich NR, Field DJ. (2012) Torosaurus Is Not Triceratops: Ontogeny in Chasmosaurine Ceratopsids as a Case Study in Dinosaur Taxonomy. PLoS ONE 7(2): e32623. doi:10.1371/journal.pone.0032623
Ostrom J.H, Wellnhofer P. (1986) The Munich specimen of Triceratops with a revision of the genus. Zitteliana. 14, 111-158.
Scannella J, Horner JR. (2010) Torosaurus is Triceratops, synonymy through ontogeny. Journal of Vertebrate Paleontology 30: 1157-1168.
Scannella, J.B. and D.W. Fowler. (2009) Anagenesis in Triceratops: evidence from a newly resolved stratigraphic framework for the Hell Creek Formation. North American Paleontological Convention , abstracts volume: 148-9.
Gregory S. Paul, Per Christiansen. (2000) Forelimb posture in neoceratopsian dinosaurs: implications for gait and locomotion Paleobiology Summer, v. 26 no. 3 p. 450-465