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Cow Shark Facts and Information:
Notorynchus - Sevengill & Hexanchus - Sixgill Cow Sharks and their Fossil Relatives.

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Sevengill Cow Shark
Sevengill Cow Shark The Sevengill Cow Shark

These series of beautiful photographs of a Sevengill Cow shark (Notorynchus cepedianus) (1) (2) (3) were taken by Derek Keats.



Fast Facts about the Cow Sharks


Types of Cow Sharks:
There are 4 species of Cow sharks, the Broadnose sevengill, Bluntnose sixgill, Sharpnose sevengill, and the Bigeyed sixgill.
The two most popular cow sharks are the Broadnose sevengill - Notorynchus cepedianus (Peron, 1807), and the Bluntnose sixgill - Hexanchus griseus (Bonnaterre, 1788).

Taxonomy:
Order: Hexanchiformes Family: Hexanchidae Genus: Notorynchus, Heptranchias, and Hexanchus Species: N. cepedianus, Hep. perlo, Hex. griseus, and Hex. nakamurai.

Age: Late Paleocene to Recent

Distribution: Global
Cow sharks have a global distribution, from tropical to temperate waters.
Sevengills can be found in shallow waters, while Sixgills are generally a deep water species.

Body Size:
Sevengills can reach sizes of around 3 m (10 feet), while the Bluntnose sixgill is longer, reaching lengths of 4.82 m (15.8 ft) (Castro, p. 27).

Diet:
The Bluntnose sixgill and Broadnose Sevengills feet on a wide variety of prey, from all sorts of bony fish and other sharks to crustaceans.

Physical Appearance:
These sharks are very primitive looking. They only have a single dorsal fin toward the back of their bodies. They are missing the characteristic large dorsal fin that is a distinctive characteristic other sharks.
The also have either six or seven gill slits, where all other sharks have five.
Their teeth are very odd looking; they are long and comb-like.

Fun Fact:
The Bluntnose sevengill sometimes hunt in packs to capture large prey!





Cow Shark Facts and Information - The Details

Goblin Shark Image

Comparison sketch of the two cow sharks, H. griseus (Broadnose sixgill), and N. cepedianus (Bluntnose sevengill) Cow Sharks.

Cow SharkOrigins, Description, and Habits

Origins of Cow Sharks
Cow sharks first appear in the fossil record in the late Cretaceous as the Hexanchus genus, while the Notorynchus genus first appears in the late Paleocene. They have not changed much in the 60 or so millino years. Today, there are four extant cow sharks. The two most common are N. cepedianus (the Bluntnose sevengill), and H. griseus (the Broadnose sixgill).


Description of Cow Sharks
Cow sharks in general have a characteristic appearance in that they only have a single dorsal fin toward the back of the shark. They are missing the large central dorsal fin that is found on most sharks. They also have very round, blunt snouts.
The sixgill sharks have (you guessed it) six gill slits, while the sevengill sharks have (right again) seven gill slits. Sixgill sharks are also larger than sevengill sharks. The Broadnose sixgill sharks have been known to reach lengths up to 4.82 m (15.8 ft) (Castro, p. 27), while sevengill sharks have been known to reach lengths of up to 2.64 m (8.7 ft) (Castro, p. 39).
The sixgill shark has a dark gray to brown color, with a lighter underside, while the sevengill shark has numerous small dark spots on the dorsal surface.


Differences and Behavior between the Cow Sharks
Although they appear similar, these sharks occupy different ecological niches. The larger Broadnose sixgill shark is primarily a nocturnal deep water shark often reported from depths of 100 - 600 fathoms (1 fathom = 6 ft = 1.8 m) (Castro, p. 27). They have a worldwide distribution in temperate and tropical waters. They feed on crustaceans and a wide array of fishes. The Bluntnose sevengill cow sharks, on the other hand, are aggressive sharks that prefer shallow near shore waters. They are often reported from depths of 3 - 6 fathoms (Castro, p. 39). They are found throughout the Indian and Pacific oceans, and feed on a variety of prey, from fish, including other sharks, to marine mammals. Bluntnose sevengills have also been observed hunting in groups, or packs, to take on larger prey.





Cow Shark Tooth Morphology - Differences/Identification between the Cow Shark Teeth


Notorynchus cepedianus tooth identification

cow shark tooth characteristics

Lower lateral Notorynchus cepedianus tooth identification diagram from Calvert Co., MD


Hexanchus griseus tooth identification

cow shark tooth characteristics

Lower lateral Hexanchus griseus tooth identification diagram from Aurora, NC.

Symphiseal Teeth

cow shark symphiseal tooth

This is a symphiseal tooth, or center tooth, from a cow shark. They are unique looking and are found in the very center of the jaw in both the upper and lower teeth.
This one was found by a nice website visitor, Jon, who found it at a quarry in Shark Tooth Hill.

Cow Shark Tooth Positions
Due to their self-evident shape, fossil Cow shark teeth are nearly impossible to confuse with other shark genera. Lower laterals have a unique rectangular shape lined with cuspls, which become smaller toward the distal end. In front of the first cusp, or primary cusp, on the mesial edge, are serrations. The roots are generally thin and rectangular shaped. Uppers are generally more square-shaped than the lowers, have less cusps, and often lack serrations on the mesial edges. Upper parasymphyseal teeth (front teeth) consist of a single angled cusp and a narrow root. Cow sharks also have a unique lower symphyseal tooth (center tooth). These teeth are fan shaped. Refer to the following image for examples of upper and lower teeth.

tooth positions of cow shark teeth

This image shows Notorynchus cepedianus upper and lower teeth from the Calvert Cliffs of MD. Notice the lower symphyseal tooth.

Hexanchus vs Notorynchus teeth - Differences
There are also some differences between male and female teeth. These differences, however, will not be discussed here.

Even though Cow shark teeth are easily differentiated from other shark genera, H. sp. and N. cepedianus can be very difficult to distinguish from each other. There are small differences between the teeth, most of which are beyond the scope of this webpage. However, here are a few general guidelines to follow. To start, H. sp. is a larger, more robust, version of N. cepedianus. The typical size of H. sp is around 32-38 mm (1.25 - 1.5 inches), while N. cepedianus is slightly smaller, around 25 mm (1 inch) (Cocke, p. 82-83). There are two key differences between the two sharks lower lateral teeth. N. cepidianus has less cusps that run behind the primary cusp. Typically there are around 5 or 6 cusps on N. cepedianus, while H. sp. has up to 13 cusps (Kent, p.18). The second difference lies in their mesial edges. H. sp. has very fine serrations on the mesial edge, while N. cepedianus has larger recurved serrations on the mesial edge. A comparison of the shoulders is shown in the image below.

hexanchus vs notorynchus cow shark teeth

This image shows a comparison between lower lateral Notorynchus and Hexanchus serrations on the mesial edge. Hexanchus has very fine serrations, while Notorynchus generally have larger, curved serrations.
This is one of the main identification differences between the two fossil shark teeth.






Fossil Examples of Cow Shark Teeth and Fossil Hunting Locations

Fossil Cow shark teeth are uncommon to find. However, they occur in many miocene and pliocene formations. Some of the more well known fossil bearing formations include: the Calvert Cliffs of Maryland and the Pungo and Yorktown formations in North Carolina.

A beautiful Hexanchus griseus fossil cow shark tooth, just exposed from the rain. Fossil Cow Shark Tooth


Hexanchus cf. collisonae, Hexanchus griseus and Notorynchus cepedianus Cow Shark Examples (Cow Sharks)




Hexanchus cf. collisonae

Eocene sixgill cow shark

This is an Eocene species of Cow Shark. These Eocene cow shark teeth use to be called Hexanchus agassizi.
Eocene Cow Shark - Virginia
Hexanchus collisonae cow shark tooth - Eocene

These fossil teeth are very fragile and hard to find. This is a lower lateral tooth.
Hexanchus collisonae is an Eocene cow shark.
Formation:Nanjemoy Formation, Potapaco Member, Bed B    Age:Lower Eocene (Ypresian), ~55 m.y.    Location:Fisher Lane Bone Bed, Fisher Branch of Muddy Creek, Staffod Co., VA    Size: ~5/8" long (15mm)   





Hexanchus griseus (Bonnaterre, 1788) aka Hexanchus gigas (Sismonda, 1857)

Broadnose sixgill cow shark

H. griseus is a living species today. The fossil teeth are probably from this extant species and first appeared in the miocene.

Hexanchus teeth can be easily differentiated from Notorynchus teeth. Notorynchus teeth have serrations on their mesial edge, whereas Hexanchus teeth have very fine serrations on the mesial edges. Also Hexanchus teeth tend to have more cones running down the length of the tooth.



1 5/8" Hexanchus (Broadnose sixgill) lower Cow Shark tooth - North Carolina
Hexanchus cow shark tooth - sixgill cow shark

This is an outstanding lower Hexanchus tooth.

Click on the image to see it when found.

Formation:Yorktown    Age:Pliocene: Roughly 2.5-5 m.y.    Location:PCS Mine, Aurora, NC    Size:~ 1 5/8" (41mm)    Date:Feb. 2009 TRIP





1 5/8" Hexanchus (Broadnose sixgill) lower Cow Shark tooth - North Carolina
Hexanchus cow shark tooth - sixgill cow shark

The teeth of Broadnose sixgill sharks are fragile and are often found broken, like the one in this image. Sadly, the conules are missing on this specimen.


Formation:Yorktown    Age:Pliocene: Roughly 2.5-5 m.y.    Location:PCS Mine, Aurora, NC    Size:~ 1 5/8" (41mm)    Date:March 2008 TRIP





Notorhynchus cepedianus (Peron, 1807) aka primigenius (Agassiz, 1843)

Bluntnose Sevengill Cow Shark

Purdy et al (2001) believe these teeth are identical to the extant N. cepedianus, and therefore N. primigenius is just synonymous to the living N. cepedianus.



1" Notorynchus Cow Shark tooth - Calvert Cliffs - Maryland
A complete Bluntnose sevengill cow shark tooth fossil from Maryland

This is my best Notorynchus tooth to date. It is nearly perfect, and was plucked straight from a clay block.
Click on the pic to see the trip this was found on.
Formation:Choptank, Zone 17?    Age:Middle Miocene ~ 15 m.y.    Location:Calvert Cliffs of MD    Size:~ 1" (25mm)    Date:June 2006 TRIP





1" Notorynchus lower Cow Shark tooth - North Carolina
Bluntnose sevengill cow shark tooth

Here is a beautiful complete Bluntnose sevengill cow shark tooth

Formation:Yorktown    Age:Pliocene: Roughly 2.5-5 m.y.    Location:PCS Mine, Aurora, NC    Size:~ 1" (25mm)    Date:Sept 2008 TRIP





1" Notorynchus lower Cow Shark tooth - North Carolina
Bluntnose sevengill cow shark tooth

Another complete Bluntnose sevengill fossil cow shark tooth
Click on the image to see it as found.
Formation:Pungo River    Age:Miocene: Roughly 18-22 m.y.    Location:PCS Mine, Aurora, NC    Size:~ 1" (25mm)    Date:Sept 2008 TRIP





1 1/8" Notorynchus Cow Shark tooth - North Carolina
A complete Bluntnose sevengill cow shark tooth fossil

Here is another complete cow shark tooth
Click on the image to see it as found
Formation:Pungo River    Age:Miocene: Roughly 18-22 m.y.    Location:PCS Mine, Aurora, NC    Size:~ 1 1/8" (27mm)    Date:March 2006 TRIP





1 1/8" Notorynchus Cow Shark tooth - North Carolina
A complete Bluntnose sevengill cow shark tooth fossil

Another complete fossil cow shark tooth

Formation:Pungo River    Age:Miocene: Roughly 18-22 m.y.    Location:PCS Mine, Aurora, NC    Size:~ 1 1/8" (27mm)    Date:Feb 2009 TRIP





Notorynchus Cow Shark teeth - Cavlert Cliffs, Maryland
A complete Bluntnose sevengill cow shark tooth fossil

This image shows different tooth positions of Notorynchus - the fossil shark teeth are incomplete

Formation:Calvert    Age:Middle Miocene: Roughly 18-15 m.y.    Location:NCalvert Cliffs of Maryland   






Recommended Books about Sharks:

Sharks of the World (Princeton Field Guides)
By: Leonard Compagno, 2005

Leonard Compagno is very thorough, so this is a VERY COMPLETE guide to sharks. This is the best shark guide I have found thus far. He catalogued all sharks from the FAO Species Catalog. There is a key to shark families, and color plates of sharks. Each individual shark has a description, drawing, examples of upper and lower teeth, distribution, size, behavior, etc...
I would recommend this book to anyone who wants to learn about the incredible diversity of sharks!




Fossil Shark Teeth of the World , A Collector's Guide
by Joe Cocke, Copyright 2002

A great book for identifying all those teeth. This book is laid out "as simple as possible." It's ease of use and small size makes it great to carry during collecting trips. This book shows teeth from around the globe, but all the Calvert teeth can be found in it.




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